中华大锹生态标本,全球深山锹物种快速一览

2022年第13期

前言:深山锹是锹甲中的大属,根据日本方面的最新统计,以及加上近年我国发布的多个新种,目前合计已有140余种深山锹(含拟深山属和太古深山属),如加上亚种(包括部分未描述的亚种),整体数量将接近170种。同时,深山锹也是我国资源最丰富的锹甲属,约有几十多种。

今天我们以国内《中华锹甲》系列著作以及日方文献(BeKuwa No.75)为基础,归纳整理了截至今年为止的全球深山锹物种列表。

分享给各位虫友,希望对喜欢深山锹的朋友有所帮助!

整个列表除深山锹属外,也包括拟深山和太古深山两个属的物种,同时深山锹按种群归纳在一起罗列,部分近年发表的新种如有原始发表文献(在线电子版),我们同样予以罗列,有需求的虫友可自行查阅。

由于内容较多,列表仅包括中文名称(如有)以及拉丁学名,不包含产地分布等其他内容,如需相关物种的详情可去我们虫社网站查询,大部分深山锹物种的资料我们已经完成收录。

一、古深山属和拟深山属

1、古深山属:Genus Noseolucanus Araya et Tanaka, 2000 

短鞘太古深山锹 Noseolucanus denticulus (Boucher, 1995)  

郑氏太古深山锹 Noseolucanus zhengi Huang, 2006

中华大锹生态标本,全球深山锹物种快速一览(1)

郑氏太古深山锹(图源:引用自文献)

2、拟深山属 :Genus Eolucanus Kurosawa, 1970 

阿迪拟深山锹 Eolucanus adi Okuda et Maeda, 2016  

大卫拟深山锹 Eolucanus davidis (Deyrolle, 1878)

贵州亚种:Eolucanus davidis guizhouensis Schenk, 2020(查阅详情)

文献:Beetles_World_22_2020

中华大锹生态标本,全球深山锹物种快速一览(2)

大卫拟深山锹贵州亚种(图源:引用自文献)

短颚拟深山锹 Eolucanus gracilis gracilis Albers, 1889  

亚种 Eolucanus gracilis sharchokpai Okuda et Maeda, 2014 

尖齿拟深山锹 Eolucanus lesnei (Planet, 1905)  

圆齿拟深山锹 Eolucanus mingyiae (Huang, 2006) 

欧氏拟深山锹 Eolucanus oberthueri (Planet, 1896) 

普氏拟深山锹 Eolucanus prometheus Boucher et Huang,1991  

潘氏拟深山Eolucanus pani Huang, 2006 

Eolucanus jonnae Schenk, 2020

文献:Beetles_World_22_2020

Eolucanus kerleyi (Boucher, 1994)

二、深山属

深山属(Genus Lucanus Scopoli, 1763):按现有分类合计有十二个种群(参考中华锹甲系列以及日本Bekuwa),其中幸运深山种群所包含的物种最多,达到了50个(不含亚种),近年国内新发表的深山锹基本都属于幸运深山锹种群;其次是藏南深山种群以及拉叉深山种群物种数量分列第二和第三。

1、幸运深山锹种群(Lucanus fortunei group)

幸运深山锹 Lucanus fortunei Saunders, 1854 

= Lucanus laevigatus Didier, 1931 (同种异名)

注:幸运深山锹为我国三有(有重要生态、科学、社会价值的陆生野生动物)昆虫

史密斯深山锹 Lucanus smithii Parry, 1862  

= Lucanus vicinus Planet, 1900 (同种异名)

黄鞘深山锹 Lucanus delavayi Fairmaire, 1887  

四川深山锹 Lucanus fairmairei Planet, 1897  

= Lucanus jietuii Nagai et Maeda, 2010 (同种异名)

越南福慕深山锹 Lucanus formosus formosus Didier, 1925 

= Lucanus cyclommatoides Didier, 1928 (同种异名:橙深山锹 ) 

未定名亚种 Lucanus formosus ssp.(越南安沛省)

雅深山锹 Lucanus nobilis Didier, 1925  

= Lucanus speciosus Didier, 1925 (同种异名,存疑)

= Lucanus cambodiensis Didier, 1925 (同种异名,存疑)

= Lucanus jeanvoinei Didier, 1927 (同种异名)

扩头深山锹 Lucanus kraatzi kraatzi Nagel, 1926  

越南亚种 Lucanus kraatzi giangae H. Ikeda, 1997  

华中深山锹 Lucanus szetschuanicus Hanus, 1932  

= Lucanus hildegardae Zilioli, 2002 (同种异名)

黄脚深山锹 Lucanus miwai Y. Kurosawa, 1966  

大明山深山锹 Lucanus deuveianus Boucher, 1998  

= Lucanus brivioi guangxii Schenk, 2009 (同种异名)

卡氏深山锹 Lucanus klapperichi Bomans, 1989  

昂山深山锹 Lucanus aungsani Zilioli, 1999 

武夷深山锹 Lucanus wuyishanensis Schenk, 1999  

均齿深山锹 Lucanus kirchneri Zilioli, 1999  

普氏深山锹 Lucanus prossi Zilioli, 2000  

= Lucanus szetschuanicus lati Nagai, 2000 (同种异名)

黄毛深山锹 Lucanus derani Nagai, 2000  

东部亚种 Lucanus derani fukinukiae Katsura et Giang, 2002(注:日方文献本种为独立物种 Lucanus fukinukiae)  

大理深山锹 Lucanus dirki Schenk, 2002 

北越深山锹 Lucanus fujitai Katsura et Giang, 2002

布氏深山锹 Lucanus brivioi brivioi Zilioli, 2003  

未定名亚种 Lucanus brivioi ssp.(布氏深山南岭亚种.广东南岭)

宽叉深山锹 Lucanus fonti Zilioli, 2005  

路克深山锹 Lucanus luci Maeda, 2010 

刘氏深山锹 Lucanus liuyei Huang et Chen, 2010  

马拉齐深山锹 Lucanus marazziorum Zilioli, 2012

中华大锹生态标本,全球深山锹物种快速一览(3)

马拉齐深山锹正模(图源:引用自文献)

钦山深山锹 Lucanus chinhillensis Schenk, 2016  

文献:Beetles World 12_2016

何氏深山锹 Lucanus hewenjiae Huang et Chen, 2013  

刘鹏宇深山锹 Lucanus liupengyui Huang et Chen, 2017 

詹氏深山锹 Lucanus zhanbishengi Wang et Zhu, 2017  

文献:Lucanus zhanbishengi sp. n., A New Species from Hunan, Central China. International Journal of Research Studies in Zoology 3(4):54-69

中华大锹生态标本,全球深山锹物种快速一览(4)

詹氏深山锹(图源:引用自文献)

朱氏深山锹 Lucanus zhuxiangi Wang et Zhan, 2018  

文献:Lucanus zhuxiangi sp. n., a New Species from Southeast China. International Journal of Research Studies in Zoology 4(3):1-11

中华大锹生态标本,全球深山锹物种快速一览(5)

朱氏深山锹(图源:引用自文献)

刘玮深山锹 Lucanus liuweii Huang et Chen, 2019 (查阅详情)  

文献:Beetles World 19_2019

岑王老山深山锹 Lucanus cenwanglaoshanus Huang et Chen, 2020 (查阅详情)

文献:Beetles World 21_2020

宇老深山锹  Lucanus yulaoensis Lin, 2021 (查阅详情)

文献:Description of Lucanus yulaoensis sp. nov., a new species stag beetle from northern Taiwan. Faunitaxys 9(2):1-5

卞氏深山锹 Lucanus shulini Bian et Zhan, 2021(查阅详情)

文献:A new species of Lucanus Scopoli, 1763 from Tibet, China. Faunitaxys 9(27):1-4.

莫氏深山锹 Lucanus moae Qi, 2021 (查阅详情)

文献:Lucanus moae sp. nov., a new species from Sichuan, China. Faunitaxys 9(23):1-7

增华深山锹 Lucanus zenghuae Wang, He, He & Zhou,2022

姬深山锹 Lucanus swinhoei swinhoei Parry, 1874  

大陆亚种 Lucanus swinhoei continentalis Zilioli, 1998  

未命名亚种 Lucanus swinhoei ssp. (中国.广西)

大屯姬深山锹 Lucanus datunensis Hashimoto, 1984  

= Lucanus ritae Lacroix, 1984 (同种异名)

Lucanus pulchellus Didier, 1925  

Lucanus bruanti Lacroix et Bomans, 1973  

Lucanus koyamai Akiyama et H. Hirasawa, 1990  

Lucanus miyashitai Mizunuma, 1994  

Lucanus pesarinii Zilioli, 1998  

Lucanus ziliolii Fukinuki, 2000  

Lucanus tsukamotoi Nagai, 2002  

= Lucanus viheari Schenk, 2013 (同种异名, 或存疑)

Lucanus adelmae Zilioli, 2003

中华大锹生态标本,全球深山锹物种快速一览(6)

Lucanus adelmae 正模(图源:引用自文献)

Lucanus satoi Nagai et Tsukamoto, 2003  

Lucanus maedai Nagai et Tsukamoto, 2003  

Lucanus takakuwai Fujita, 2010  

Lucanus jibii Okuda et Maeda, 2015  

Lucanus gradivus Sato et Zilioli, 2017

文献:Lucanus gradivus n. sp. from Vietnam, with new records of L. fujitai Katsura & Giang from Vietnam and Laos. Natural History Sciences. Atti Della Società Italiana Di Scienze Naturali E Del Museo Civico Di Storia Naturale Di Milano 5(1):11-15.

中华大锹生态标本,全球深山锹物种快速一览(7)

Lucanus gradivus(图源:引用自文献)

2、藏南深山锹种群(Lucanus lunifer group,注)

注:日方资料将本种群列为 Lucanus villosus 种群,我们这里参考《中华锹甲》列为藏南深山种群。

短顎深山锹 Lucanus imitator Boucher et Huang, 1991  

= Puseudolucanus bicolor Schenk, 1996 (同种异名)

郎氏深山锹 Lucanus langi Huang, He et Shi, 2011  

文献:Description of two new species of Lucanus Scopoli from southeastern Tibet, China. Zootaxa 2987:56-64.

中华大锹生态标本,全球深山锹物种快速一览(8)

郎氏深山锹(图源:引用自文献)

周氏深山锹 Lucanus choui Huang et Chen, 2013  

陈氏深山锹 Lucanus cheni Huang, He et Shi, 2011  

= Lucanus furcifer Arrow, 1950 (同种异名:佛深山锹)

文献:Description of two new species of Lucanus Scopoli from southeastern Tibet, China. Zootaxa 2987:56-64.

藏深山锹 Lucanus thibetanus tibetanus Planet, 1898    

独龙江亚种 Lucanus thibetanus isaki Nagai, 2000  

怒江亚种 Lucanus thibetanus singularis Planet, 1900  

= Lucanus gennestieri Lacroix, 1971 (同种异名 )

云南亚种 Lucanus thibetanus furcifer Arrow, 1950  

= Lucanus pseudosingularis Didier er Seguy, 1953 (同种异名 )

= Lucanus thibetanus katsurai Mizunuma, 1994 (同种异名 )

未定名亚种 Lucanus thibetanus ssp (梵浄山亚种)

藏南深山锹 Lucanus lunifer lunifer Hope, 1833  

= Lucanus rugifrons Hope et Westwood, 1845 (同种异名 )

东部亚种 Lucanus lunifer franciscae Lacroix, 1971 

盈江深山锹 Lucanus victorius Zilioli, 2002  

弗瑞深山锹 Lucanus fryi Boileau, 1911  

= Lucanus fryi schepanski Schenk, 2008 (同种异名)

Lucanus villosus Hope, 1831  

Lucanus westermanni Hope et Westwood, 1845  

Lucanus izindui Okuda et Maeda, 2016  

拉叉深山锹种群(Lucanus laminifer group)

拉叉深山锹 Lucanus laminifer laminifer Waterhouse, 1890  

= Lucanus laminifer coronatus Zilioli, 2000 (同种异名)

= Lucanus laminifer lucidulus Zilioli, 2000 (同种异名)

亚种 Lucanus laminifer kusakabei Fujita, 2010  

台湾深山锹 Lucanus formosanus Planet, 1899   

巨叉深山锹 Lucanus hermani De Lisle, 1973(注:本种为国家二级保护野生动物) 

普叉深山锹 Lucanus planeti planeti Planet, 1899  

广西亚种 Lucanus planeti dayaoshanensis Schenk, 2011  

= Lucanus planeti guangxii Schenk, 2009 (同种异名)

维氏深山锹/维叉深山锹 Lucanus vitalisi Pouillaude, 1913   

牛叉深山锹 Lucanus niu Wang, He, He & Zhou,2022

中华大锹生态标本,全球深山锹物种快速一览(9)

牛叉深山锹正模(图源:引用自文献)

三、其他深山锹种群

1、日本深山锹种群(Lucanus maculifemoratus group)

日本深山锹 Lucanus maculifemoratus maculifemoratus Motschulsky, 1861  

伊豆深山锹(日本深山锹伊豆亚种) Lucanus maculifemoratus adachii Tsukawaki, 1995 

斑股深山锹 Lucanus dybowskyi dybowskyi Parry, 1873  

= Lucanus dybowskyi jilinensis Li, 1992 (同种异名)

台湾亚种 Lucanus dybowskyi taiwanus Miwa, 1936  

四川亚种 Lucanus dybowskyi lhasaensis Schenk, 2006 

黑泽深山锹 Lucanus kurosawai Sakaino, 1995

栗色深山锹 Lucanus kanoi kanoi Y. Kurosawa,1966  

北部亚种 Lucanus kanoi piceus Y. Kurosawa, 1966 

东南亚种(黑脚深山锹) Lucanus ogakii ogakii Imanishi, 1990  

=Lucanus ogakii chuyunsanus Sakaino et Yu, 1993(同种异名:黑脚深山锹出云山亚种 )

注:日方文献将黑脚深山锹作为独立物种,并仍有”出云山亚种“。 

梵净深山锹 Lucanus fanjingshanus Huang et Chen, 2010  

Lucanus takeoi Adachi, 2020  

2、瑟深山锹种群(Lucanus sericeus group)

瑟深山锹 Lucanus sericeus sericeus Didier, 1925  

亚种 Lucanus sericeus teshii Nagai et Tsukamoto, 2003

亚种 Lucanus sericeus ohbayashii Nagai et Tsukamoto, 2003

南山深山锹 Lucanus nangsarae Nagai, 2000 

3、金属深山锹种群(Lucanus mearesii group)

金属深山锹 Lucanus mearesii Hope, 1842  

鬼深山锹 Lucanus wemckeni Schenk, 2006  

Lucanus rondoi Okuda et Maeda, 2018 

4、康拓深山锹种群(Lucanus cantori group)

康拓深山锹 Lucanus cantori cantori Hope, 1842  

亚种 Lucanus cantori colasi Lacroix, 1967

简颚深山锹 Lucanus didieri Planet, 1927  

Lucanus groulti (Planet, 1897) 注:存在不明种

5、弯叉深山锹种群(Lucanus angusticornis group)

弯叉深山锹 Lucanus angusticornis Didier, 1925  

= Lucanus angusticornis inclinatus Schenk, 2008 弯叉深山锹中国亚种 (同种异名或存疑)

越南深山锹 Lucanus ngheanus ngheanus Okuda, 2009  

昆嵩亚种 Lucanus ngheanus phuongi Okuda et Maeda, 2012 

兔儿深山锹 Lucanus dongi Maeda, 2009  

奄美深山锹 Lucanus ferriei Planet, 1898  

6、派瑞深山锹种群(Lucanus parryi group)

帕瑞(派瑞)深山锹 Lucanus parryi Boileau, 1899 

西部亚种 Lucanus parryi laetus Arrow, 1943(注) 

中华大锹生态标本,全球深山锹物种快速一览(10)

帕瑞(派瑞)深山锹

中华大锹生态标本,全球深山锹物种快速一览(11)

帕瑞(派瑞)深山锹

注:日本方面认为派瑞西部亚种为独立种,既Lucanus laetus。

御藏深山锹 Lucanus gamunus Sawada et Watanabe, 1960  

承远深山锹 Lucanus chengyuani Wang et Ko, 2018(查阅详情) 

7、黄胫深山锹种群(Lucanus boileaui group)

黄胫深山锹 Lucanus boileaui Planet, 1897  

= Lucanus bidentis Schenk, 2013 (同种异名)

路氏深山锹 Lucanus ludivinae Boucher, 1998  

8、Lucanus manai 种群(Lucanus manai group)

Lucanus manai Bomans et Miyashita, 1997  

Lucanus nosei Nagai, 2000  

= Lucanus werneri Zilioli, 2000 (同种异名)

双齿深山锹 Lucanus nyishwini nyishwini Nagai, 2000  

墨脱亚种 Lucanus nyishwini bretschneideri Schenk, 2008 

9、Lucanus dohertyi 深山锹种群(Lucanus dohertyi group)

Lucanus dohertyi Boileau, 1911  

Lucanus sukkiti Fukinuki, 2004 注:存在不明种(尼泊尔等)

直颚深山锹 Lucanus hayashii Nagai, 2000  

= Lucanus hansi Zilioli, 2000 (同种异名)

米什米深山锹 Lucanus mishmi Okuda et Maeda, 2014 

四、欧美深山属物种

1、欧洲等地区深山锹

阿深山锹 Lucanus atratus Hope, 1831 注:我国西藏也有分布 

Lucanus confuses (Boucher, 1994)   

Lucanus wittmeri (Lacroix, 1984)  

Lucanus deuvei deuvei (Lacroix, 1988)  

亚种 Lucanus deuvei pakistanicus Schenk, 2009  

Lucanus feglini Lacroix et Bomans, 1973  

薛西斯深山锹 Lucanus xerxes Kral, 2004 

文献:Lucanus (Pseudolucanus) xerxes sp. nov. from the Zagros Mts., with some distribution data to the lucanid fauna from Iran. Acta Societatis Zoologicae Bohemicae 68(3):183-189.

Lucanus busignyi (Planet, 1909)  

Lucanus syriacus Motschulsky, 1870  

Lucanus macrophyllus Kraatz, 1860  

巴巴罗萨深山锹 Lucanus barbarossa (Fabricius, 1801)  

伊比利亚深山锹 Lucanus ibericus Motschulsky, 1845  

Lucanus subvelutinus Motschulsky, 1870 注:存在多个不明种,部分学者认为本种为伊比利亚深山锹亚种

中华大锹生态标本,全球深山锹物种快速一览(12)

24-25 Lucanus ibericus 26-27 Lucanus subvelutinus(图源:引用自文献)

Lucanus tetraodon tetraodon Thunberg, 1806  

亚种 Lucanus tetraodon provincialis Colas, 1949  

亚种 Lucanus tetraodon sicilianus Planet, 1899 

欧洲深山锹 Lucanus cervus cervus (Linne, 1758)  

阿卡巴深山锹(欧洲深山锹阿卡巴亚种) Lucanus cervus akbesianus Planet, 1896 

犹太深山锹(欧洲深山锹犹太亚种) Lucanus cervus judaicus Planet, 1902  

土耳其深山锹(欧洲深山锹土耳其亚种) Lucanus cervus turcicus Sturm, 1843  

Lucanus cervus poujadei Planet, 1897 

Lucanus pontbrianti (Mulsant, 1839)  

Lucanus laticornis Deyrolle, 1864  

菲德勒深山锹  Lucanus fiedleri Schenk et Fiedler, 2020  

文献:Beetles World 21_2020

中华大锹生态标本,全球深山锹物种快速一览(13)

菲德勒深山锹(图源:引用自文献)

2、美洲地区深山锹

美国深山锹 Lucanus mazama (LeConte, 1861)  

红鹿深山锹 Lucanus elaphus Fabricius, 1775 

中华大锹生态标本,全球深山锹物种快速一览(14)

红鹿深山锹生态照(图源:引用自文献)

Lucanus capreolus (Linne, 1763)  

Lucanus placidus Say, 1825 

后记:目前仍旧有很多未描述的深山锹物种(亚种),不排除未来继续有新种发表;同时部分深山锹分类上存在一定问题或争议,比如欧洲地区的物种,以上分类不排除后续有新调整,当前列表也仅供参考!

参考文献:

1、中华锹甲1-3:

2、A checklist of stag beetles (Coleoptera: Scarabaeoidea: Lucanidae) from Iran November 2014Zootaxa 3887(3):422-436

3、Beetles World 12、19、21、22

4、A review of geographical distribution of the stag beetles in Mediterranean countries Fragmenta entomologica, 48 (2): 153-168 (2016)

5、New descriptions and notes on Chinese stag-beetles, with discovery of the second species of Noseolucanus from SE Tibet.

6、中国锹甲科四新记录种记述(鞘翅目:锹甲科:锹甲亚科)

以上如有错误还望指正,感谢阅读!

注:欢迎转载,标明出处即可!

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